The ichnological survey of the track-bearing surface in the Pontrelli quarry (Altamura, Apulia) was performed using both traditional and new methodologies. Approximately 26,000 footprints were identified, 600 of which arranged in 12 quadrupedal trackways. Close-range photogrammetry was applied to gain accurate 3D models of the best-preserved specimens. The ichnological study, supported by morphometric analysis and photogrammetric models, led to the identification of different morphotypes, produced by two dinosaur trackmakers. In the Morphotype 1 are included the specimens characterised by tridactyl pes, which digits are lobe-shaped, and smaller manus (i.e., heteropody index between 1:5 and 1:2) with three subcircular pads. This ichnotaxonomical group includes the ichnospecies Apulosauripus federicianus, established by Nicosia et al. (2000) to describe the trackway ACDL99/3 . Nevertheless, numerous footprints can be also morphologically matched to the ichnogenus Caririchnium Leonardi, 1984. Six trackways can be assigned to this morphotype, including the two longest trackways of the ichnosite (represented by 75 and 88 manus-pes couples). The specimens of Morphotype 1 are referred to ornithopod dinosaurs, likely to medium-sized hadrosaurs with a hip height between 74 and 125 cm and up to 4.5 m long. The closest candidate is to date Tethyshadros insularis, 4-5 m long hadrosauroid from the upper Campanian-lower Maastrichtian deposits of Villaggio del Pescatore (Trieste, NE Italy). The Morphotype 2 is represented by a broad tetradactyl pes and a slightly smaller manus (i.e., heteropody index between 1:3 and 1:2 on average, in some specimens up to 1:1), in which four or five digits are barely recognizable. Six trackways are referable to this morphotype, among which the longest is constituted by 39 quadrupedal steps. The morphological features of these tracks closely match those of to the ichnogenus Tetrapodosaurus Sternberg, 1932, typically attributed to ankylosaurs. According primarily to the medium size of the ankylosaurian trackmaker of Altamura (hip height = 60-92 cm, body length = 2.5-4.5 m), the coincidence correlation method (Carrano & Wilson, 2001) suggests a nodosaurid as most suitable candidate, similar to the 2.5 m long Struthiosaurus or the 4 m long Hungarosaurus from the Santonian-Maastrichtian deposits of Europe. The occurrence of these two dinosaur groups in the Late Cretaceous of the Apulian Carbonate Platform can represent the keystone to enhance the knowledge on the palaeogeographic framework of the Periadriatic region.

The hadrosaur and nodosaur ichnoassemblage from the Altamura dinosaur tracksite (early Campanian; Apulia, southern Italy)

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The hadrosaur and nodosaur ichnoassemblage from the Altamura dinosaur tracksite (early Campanian; Apulia, southern Italy)

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