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Infrared unmanned aerial vehicle (UAV) target images often suffer from motion blur degradation caused by rapid sensor movement, significantly reducing contrast between target and background. Generally, detection performance heavily depends on the discriminative feature representation between target and background. Existing methods typically treat deblurring as a preprocessing step focused on visual quality, while neglecting the enhancement of task-relevant features crucial for detection. Improving feature representation for detection under blur conditions remains challenging. In this paper, we propose a novel $\textbf{J}$oint $\textbf{F}$eature-$\textbf{D}$omain $\textbf{D}$eblurring and $\textbf{D}$etection end-to-end framework, dubbed $JFD^{3}$. We design a dual-branch architecture with shared weights, where the clear branch guides the blurred branch to enhance discriminative feature representation. Specifically, we first introduce a lightweight feature restoration network, where features from the clear branch serve as feature-level supervision to guide the blurred branch, thereby enhancing its distinctive capability for detection. We then propose a frequency structure guidance module that refines the structure prior from the restoration network and integrates it into shallow detection layers to enrich target structural information. Finally, a feature consistency self-supervised loss is imposed between the dual-branch detection backbones, driving the blurred branch to approximate the feature representations of the clear one. We also construct a benchmark, named IRBlurUAV, containing 30,000 simulated and 4,118 real infrared UAV target images with diverse motion blur. Extensive experiments on IRBlurUAV demonstrate that $JFD\^{3}$ achieves superior detection performance while maintaining real-time efficiency. Code and dataset are released at \textit{https://anonymous.4open.science/r/JFD3-76C1}.